Grizzly 399 became a celebrity of Grand Teton National Park in her lifetime. Now, her death has drawn attention to wildlife-vehicle collisions and how they might be reduced

The death of grizzly No. 399 saddened tourists, biologists and photographers who for years captured images of the bear and her many cubs.

The driver who struck and killed a famous grizzly bear south of Grand Teton National Park earlier this week was not speeding and the bear’s death was an accident, law enforcement officers said Friday.